Ph.D., Population Biology, UC Davis
Dissertation: "Ecological factors affecting tetraploid speciation in snow buttercups (Ranunculus adoneus: Ranunculaceae)."
Advisor: Dr. Maureen L. Stanton
M.A., Secondary Science Education, Lewis and Clark College, Portland, OR
B.A., summa cum laude, Philosophy, Carleton College, Northfield, MN